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玦 i i i 寥了任遗世 寥我耪臀宋美 . . 茵屙赍眸, ...

-- [ 羊疣龛鲟 2 ] --

= (tf t0 ) Pm (2.4) 填戾眚 怅膻麇龛 镳铗铌 topt, 铒疱溴栝 爨犟桁嚯 箴铈嚅 镳 翳犟桊钼囗眍, 祛骓 i i i i i i i i 物蜩扈玎鲨 怵屐屙 怅膻麇龛 镳铗铌 箐咫铋 耜铕铖蜩 磬轵 桤 篑腩忤 疣忮眈蜮 眢膻 镳铊玮钿眍 镱 t 恹疣驽龛 (2.4). 项漶蜞怆 恹疣驽龛 (2.5) B0 = Pm (t0 tl ) + Bl, 恹疣驽龛 (2.4) Pm Y = (tf t0 ) Pm Pm (t0 tl ) Bl [1 e(tf t0 ) ] 滂趑屦屙鲨痼 镱 t0, 镱塍麒 dY = Pm (Pm ) [1 e(tf t0 ) ]+ dt Pm Pm (t0 tl ) Bl e(tf t0 ) = + = Pm + Pm Pm e(tf t0 ) + + (Pm Pm (t0 tl ) Bl ) e(tf t0 ).

磬觐礤, dY = [ Pm (t0 tl ) Bl ] e (tf t0 ) = 0.

dt 悟颃溧 Bl t0 = topt = tl Pm i i i i i i i i 70 茵屙赍眸 . ., 缅忸疸桤 . ., 令痤怅钼 . .

项耜铍 topt tl, 耥, 黩 铒蜩爨朦眍 珥圜屙桢 祛戾眚 怅膻麇龛 镳铗铌 脲骅 玎 镳邃咫囔 腓礤 眍 羿琨 痤耱 (痂. 2.2). 袜 腓礤轫铎 篦囫蜿 磬觐 镨蝈朦眍 牮桠铋 趔黻鲨 Y = Y (t0 ) 筢噱. 央邃 忄蝈朦眍, 祛戾眚 怅膻麇龛 镳铗铌, 耦铗忮蝰蜮簋 爨犟桁嚯铎 箴铈帼, 耦怙噤噱 祛戾眚铎 磬鬣豚 腓礤轫钽 痤耱. 青戾蜩, 蝾麝 磬鬣豚 腓礤轫钽 痤 耱 踵疣牝屦桤箦蝰 爨犟桁嚯铋 镳钿箨蜩忭铖螯 爨犟桁嚯铋 箐咫铋 耜铕铖螯 痤耱 扈牮钼钿铕铖 脲.

洛腓麒睇 B0 恹疣驽龛 (2.4) 铒疱溴脲睇 B l, . . 钽疣龛麇睇 扈龛 镳铎彐篁赅 Bl B 爨朦眍 爨犟桁嚯铋 镫铗眍耱 牦朦蝮瘥 疣 赅 腓礤轫钽 痤耱.

杨铗忮蝰蜮屙眍, 箐咫 耜铕铖螯 镳铗铌 狍溴 钽疣龛麇磬 镳邃咫囔 Pm Pm, max min, Bl Bl 黩 溧蛤 麇螓疱 玎戾鬣蝈朦睇 耠篦 恹犷疱 疱 骅爨 牦朦蜩忤痤忄龛 疣珉梓睇扈 箐咫铋 耜铕 耱 镳铗铌 磬鬣朦眍 徼铎囫耦, 溻 桤 觐蝾瘥 耱圉桀磬痦 滂磬扈麇耜桢 耦耱龛 (痂. 2.3). 橡 爨犟桁嚯铋 箐咫铋 耜铕铖蜩 镳铗铌 Pm max = Bl 镳钼邃胳 耩噔礤龛 戾驿 溻箪 耠篦扈: 1) 镳铗铌 怅膻鞲 磬鬣脲 腓礤轫钽 痤耱 tl ;

2) 镳铗铌 怅膻鞲 i i i i i i i i 物蜩扈玎鲨 怵屐屙 怅膻麇龛 镳铗铌 箐咫铋 耜铕铖蜩 觐眦 腓礤轫钽 痤耱 tl. 蔓疣驽龛 (2.4) 镳桧桁噱 耦铗忮蝰蜮簋 溧眄 耠篦 忤:

Pm (2.6) Y t0 = tl, = = (tf tl ) Pm ;

Bl Pm t0 = tl, = = (tf tl ) Pm + Y Bl l + (B l Bl ) 1 ePm /Bl (tf t ). (2.7) 骂 怛铕铎 耠篦噱 祛戾眚 怅膻麇龛 镳铗铌 tl 箴铈嚅 潆 镥疴钽 耦耱噔栩 (tl tl ) Pm (. . 箴铈嚅 玎 镳铎彐篁铌 怵屐屙 [tl, tl ]). 如 (2.7) 耠邃箦, 黩 筲咫梓屙桢 怵屐屙 牦朦蜩忤痤忄龛 疣珥桷 箴 驵 狍溴 箪屙囹. 镳 tf 箴铈嚅 狍溴 钿桧嚓钼 潆 钺铊 耠篦噱 (痂. 2.4).

理嚯钽梓眍 耩噔礤龛 祛骓 镳钼羼蜩 潆 tl tl 镳 扈龛爨朦眍 箐咫铋 耜铕铖蜩 镳铗铌 Pm min =, Bl 觐蝾痤 蜞赕 镱赅驽 鲥脲耦钺疣珥铖螯 怅膻麇龛 镳铗铌 磬鬣脲 腓礤轫钽 痤耱 (痂. 2.5) Pm Y t0 = tl, = = (tf tl ) Pm Bl l (B l Bl ) 1 ePm /B (tf tl ), (2.8) i i i i i i i i 72 茵屙赍眸 . ., 缅忸疸桤 . ., 令痤怅钼 . .

需. 2.2. 青忤耔祛耱 忮腓麒睇 箴铈 铗 祛戾眚 怅膻麇龛 镳铗铌.

i i i i i i i i 物蜩扈玎鲨 怵屐屙 怅膻麇龛 镳铗铌 箐咫铋 耜铕铖蜩 需. 2.3. 义铕弪梓羼觇 牮桠 滂磬扈觇 镫铗眍耱 (B) 牦朦蝮瘥 Dunaliella salina Teod. 潆 爨犟桁嚯铋 扈龛爨朦眍 耜铕铖蝈 镳铗铌, 潆 溻篚 祛戾眚钼 怅膻麇龛 镳铗铌 磬鬣脲 觐眦 腓礤轫钽 痤耱.

i i i i i i i i 74 茵屙赍眸 . ., 缅忸疸桤 . ., 令痤怅钼 . .

需. 2.4. 佯噔龛蝈朦磬 铞屙赅 忮腓麒睇 箴铈 镳 爨犟桁嚯铋 箐咫铋 耜铕铖蜩 镳铗铌: 镳铗铌 怅膻 鞲 磬鬣脲 (1) 觐眦 (2) 腓礤轫钽 痤耱.

i i i i i i i i 物蜩扈玎鲨 怵屐屙 怅膻麇龛 镳铗铌 箐咫铋 耜铕铖蜩 需. 2.5. 佯噔龛蝈朦磬 铞屙赅 忮腓麒睇 箴铈 镳 扈龛爨朦眍 箐咫铋 耜铕铖蜩 镳铗铌: 镳铗铌 怅膻 鞲 磬鬣脲 (1) 觐眦 (2) 腓礤轫钽 痤耱. Yl 箴铈嚅 磬 祛戾眚 tl.

i i i i i i i i 76 茵屙赍眸 . ., 缅忸疸桤 . ., 令痤怅钼 . .

Pm t0 = tl, = = (tf tl ) Pm. (2.9) Y Bl 篷腓 徼铎囫耋 B, 磬躅 牦朦蜩忄蝾疱 磬 祛戾眚 怵屐屙 tf, 铗礤耱 蜞赕 箴铈帼, 蝾 Pm B0 穂1e(tf t0 ) ]+ Y +B = (tf t0 )稰m Pm Pm B0 積(tf t0 ) = (tf t0 )稰m +B0.

+ 予螓忄 (2.5), 箴铈嚅, 铟邂桎眍, 狍溴 疣忮 Pm (tf tl ) + Bl.

橡桁屙屙桢 祛溴腓. 袜 痂. 2.3 镳邃耱噔脲磬 囡 镳铌耔爨鲨 耧屦桁屙蜞朦睇 溧眄 腓礤轫铋 羿 琨 磬觐镨蝈朦眍 牮桠铋 疣聍蛤 滂磬扈觇 镫铗眍耱 牦朦蝮瘥 D. salina 镳 疣珉梓睇 箐咫 耜铕铖 镳铗铌 祛戾眚圊 邈 怅膻麇龛. 项漶麒蜞屐, 赅 觐 箴铈嚅 祛骓 镱塍麒螯 怛鬻屙桢 钿眍泐 戾鲟 镳 爨犟桁嚯铋 扈龛爨朦眍 箐咫 耜铕铖 镳铗铌, 铒桊顸 磬 疣聍蛤睇 溧眄 腓礤轫铋 羿琨 磬觐镨蝈朦眍 牮桠铋 牦朦蝮瘥 D. salina. 项漶蜞忤 溧眄 桤 蜞犭桷 2.2 恹疣驽龛 (2.6), (2.7), (2.8) (2.9).

橡 爨犟桁嚯铋 箐咫铋 耜铕铖蜩 镳铗铌 3, = 0,19 耋 max = 18, i i i i i i i i 物蜩扈玎鲨 怵屐屙 怅膻麇龛 镳铗铌 箐咫铋 耜铕铖蜩 镱塍麒 镥疴铎 耠篦噱 (镳铗铌 怅膻鞲 磬鬣脲 腓 礤轫钽 痤耱 祛戾眚 怵屐屙 tl ):

Yl,=max = (30 0) 3,59 = 107,7 2 ;

忸 怛铕铎 (镳铗铌 怅膻鞲 磬鬣脲 腓礤轫钽 痤耱 祛戾眚 tl ):

l Y=max =(30 7) 3,59+ +(44,08 18,85)(1 e0,19 (307) ) = 107,48 惴2.

亦犭桷 2.2: 泥眄 磬觐镨蝈朦眍 牮桠铋 袜桁屙钼囗桢 琼圜屙桢 朽珈屦眍耱 2 否篁 3, 橡钿箨蜩忭铖螯, Pm 想铗眍耱 牦朦蝮瘥 磬- 18, 鬣脲 腓礤轫钽 痤耱, Bl 想铗眍耱 牦朦蝮瘥 44, 觐眦 腓礤轫钽 痤耱, B l 袜鬣腩 腓礤轫铋 羿琨 痤- 0 耋 耱, tl 暑礤 腓礤轫铋 羿琨 痤- 7 耋 耱, tl 锑犟桁嚯 箐咫 耋 0, 耜铕铖螯 镳铗铌, max 惕龛爨朦磬 箐咫 耋 0, 耜铕铖螯 镳铗铌, min i i i i i i i i 78 茵屙赍眸 . ., 缅忸疸桤 . ., 令痤怅钼 . .

西邂桎眍, 珥圜屙 忮腓麒睇 箴铈 潆 钺铊 耠篦噱 犭桤觇. 朽珉梓 狍潴 蝈 犷朦, 麇 戾睃  怵屐 牦朦蜩忤痤忄龛 (痂. 2.6).

理嚯钽梓眍 潆 扈龛爨朦眍 忮腓麒睇 min :

3, = 0,08 耋1, min = 44, 镳 怅膻麇龛 镳铗铌 祛戾眚 磬鬣豚 腓礤轫钽 痤 耱 tl 箴铈嚅 耦耱噔栩 Yl,=min =(30 0) 3, (44,0818,85) (1 e0,08(300) ) = 84,76 2.

镳 怅膻麇龛 镳铗铌 觐眦 腓礤轫钽 痤耱 l Y=min = 23 3,59 = 82,57 2.

亦觇 钺疣珙, 磬 铖眍忮 恹桤腩驽眄钽 祛 眍 玎觌栩: 麇 犷朦 怵屐 牦朦蜩忤痤忄龛 ( 疣礻圊 腓礤轫钽 痤耱 蹂祛耱囹), 蝈 戾睃 忮 腓麒磬 箴铈 玎忤耔 铗 祛戾眚 怅膻麇龛 镳铗铌, 镫铗眍耱 牦朦蝮瘥 箐咫铋 耜铕铖蜩 镳铗铌.

橡邃腩驽眄 祛溴朦 篑腩忤 镳铎脲眄钽 镳铊玮钿耱忄 徼铎囫覃 D. salina 镱玮铍桦 疣琊噌铗囹 耠邃簋 疱觐戾礓圉梃: 潆 镱塍麇龛 爨犟桁嚯 泐 箴铈 镳 钽疣龛麇眄铎 怵屐屙 牦朦蜩忤痤忄龛 礤钺躅滂祛 铕汔龛珙忄螯 蹂祛耱囹 1) 爨犟桁嚯铋 忮腓麒眍 箐咫铋 耜铕铖蜩 镳铗铌 ( 疣礻圊 腓礤 眍泐 痤耱), 2) 怅膻麇龛屐 镳铗铌 磬鬣脲 腓礤 眍 羿琨 痤耱.

i i i i i i i i 蔫磬扈赅 觐眦屙蝠圉梃 NaCl 牦朦蝮疣朦眍 耩邃 需. 2.6. 青忤耔祛耱 忮腓麒睇 箴铈 D. salina (BY ) 铗 怵屐屙 牦朦蜩忤痤忄龛 镳 = 0,19 耋1 : 1 镳铗铌 怅膻鞲 祛戾眚 怵屐屙 tl, 2 镳铗铌 怅膻 鞲 祛戾眚 怵屐屙 tl.

i i i i i i i i 80 茵屙赍眸 . ., 缅忸疸桤 . ., 令痤怅钼 . .

2.2. 蔫磬扈赅 觐眦屙蝠圉梃 NaCl 牦朦蝮疣朦眍 耩邃 橡 恹疣忄龛 Dunaliella salina 鲥朦 镱塍麇龛 -赅痤蜩磬 觐眦屙蝠圉 NaCl 镨蜞蝈朦眍 耩邃 怆弪 箫疣怆桁 羿牝铕铎 赅痤蜩眍沐礤玎 [14].

禹桕嚯 铖钺屙眍耱 汔腩犴 扈牮钼钿铕铖脲 恹 溴疰桠囹 镱恹眄簋 耦敫眍耱 耧铖钺耱忸忄豚 疣 疣犷蜿 蝈蹴铍钽栝 恹疣忄龛 D. salina 牦朦蜩忄 蝾疣 铗牮钽 蜩镟 (铉屦圊, 徉耨彘磬), 沅 恹耦觇 觐眦屙蝠圉梃 NaCl 钿眍 耱铕铐 恹琨忄 桧蝈 耔忭 耔眚彗 赅痤蜩眍, 漯筱铋 怆 觌 恹 祛戾眚铎 镱滗屦驵龛 嚯铍钽梓羼觐 麒耱 螓 牦朦蝮瘥. 唏眍, 黩 蜞觇 篑腩忤 礤镳屦眍 牦朦蜩忤痤忄龛 蝠遽箦 珥圜栩咫钽 疣聃钿 耦脲 疋玷 桉镱朦珙忄龛屐 镨蜞蝈朦睇 耩邃 钽痤祉铋 爨耨钼铋 漕脲 NaCl (耩邃 鲤蜞痂 116 /, Ben-Amotz 60 / 漯.). 泥驽 镳 扈龛爨朦睇 钺爨 镳铊 忸漶蜮 疣聃钿 NaCl 桉麒耠弪 蝾眄囔.

酿筱 礤 戾礤 忄骓 镳钺脲爨 镳铊玮钿耱忄 徼铎囫覃 D. salina 玎觌屙 耧铖钺 铗溴脲龛 徼 爨耨 铗 耩邃 镳 襻铕 箴铈. 武龛 桤 襦禧 镳铖螓 疣耧痤耱疣礤眄 怆弪 耧铖钺 铖噫溴 龛 觌弪铌 鲥眚痂趔汨痤忄龛屐. 武磬觐 镳 桉镱朦珙 忄龛 钺睇 鲥眚痂趔 磬栳铍邋 溴忸泐 钺铕 漕忄龛 镳铊玮钿耱忮眄 爨聒蜞徉 蜞觐 耧铖钺 疋玎 犷朦扈 怵屐屙睇扈 玎蝠囹囔. 项铎 磬 铎 囡 耋耱忮眄簋 痤朦 桡疣弪 钺 鲥眚痂 i i i i i i i i 蔫磬扈赅 觐眦屙蝠圉梃 NaCl 牦朦蝮疣朦眍 耩邃 趔汨痼屐铋 耋耧屙玷. 族 戾睃 铗 钺, 蝈 戾睃 怵屐屙 镱蝠遽箦 囡 皴镟痂痤忄龛.

鲥朦 疱泱鲨 觐眦屙蝠圉梃 NaCl 牦朦蜩 忄蝾疱 镱耩邃耱忸 桤戾礤龛 箐咫铋 耜铕铖蜩 镳 蝾赅 耩邃 箪屙屙 钺爨 鲥眚痂趔汨痼屐铋 耋耧屙玷 徨 镱蝈痂 徼铎囫覃 磬 囡 襻铕 箴铈 猁豚 疣琊噌铗囗 镳铖蝈轼 爨蝈爨蜩麇耜 祛溴朦 铒蜩扈玎鲨 蝈蹴铍钽梓羼觐泐 镳铞羼襦 恹疣忄龛 D. salina 牦朦蜩忄蝾疣 铗牮钽 蜩镟.

项耱囗钼赅 玎溧麒. 象耱 桁邋蝰 牦朦蜩忄蝾 铗牮 蝾泐 蜩镟 (徉耨彘) 疣犷麒 钺祛 W 耋耧屙 玷彘 扈牮钼钿铕铖脲, 恹疣忄屐 礤镳屦眍 疱骅戾. 青 聍蛤 桉镟疱龛 忸潲, 桉觌 镳铞羼覃 钺疣珙忄龛 铍疙钽 镟疣, 镱忮瘐眍耱 耋耧屙玷 牦朦蜩忄蝾疱 忮腓麒睇 箐咫铋 耜铕铖蜩 镳铗铌 磬 怩钿 磬 恹躅溴 牦朦蜩忄蝾疣 礤钿桧嚓钼. 象耱 蝈 牦 觐眦屙蝠圉 NaCl 徉耨彘礤 S0, 觐眦屙蝠 鲨 桉镱朦珞屐铋 镨蜞蝈朦眍 耩邃 S. 橡邃镱豚汔, 黩 玎溧眄 觐祜铐屙 镨蜞蝈朦眍 耩邃 礤 镱蝠遽 弪 扈牮钼钿铕铖扈, 礤钺躅滂祛 磬轵 趔黻鲨, 铒桉帼 滂磬扈牦 觐眦屙蝠圉梃 NaCl 耋耧屙 玷 潆 蹂祛耱囹 赈噻桧屣疱瘥忭铋 牦朦蝮瘥.

填溴朦. 湾镳屦睇 疱骅 牦朦蜩忤痤忄龛. 朽 耢铗痂 牦朦蝮痼 扈牮钼钿铕铖脲 蹂祛耱囹 箐咫 眍 耜铕铖螯 镳铗铌 镳 箐咫铋 耜铕铖螯 桉 镟疱龛 忸潲 桉. 亦赕 忖邃胳 疣耨祛蝠屙桢 耠 潴桢 忮腓麒睇: m0 蝈牦 觐腓麇耱忸 耦腓 耋耧屙玷, ;

m恹 觐腓麇耱忸 耦腓 恹眍耔祛 i i i i i i i i 82 茵屙赍眸 . ., 缅忸疸桤 . ., 令痤怅钼 . .

镳铗铌铎, ;

m镳 觐腓麇耱忸 耦腓 忭铖桁铄 镳铗 觐 牦朦蜩忄蝾, ;

v镳 耜铕铖螯 镱溧麒 镨蜞蝈朦 眍 耩邃 牦朦蜩忄蝾, /耋;

v恹 耜铕铖螯 恹 蝈赅龛 耋耧屙玷 桤 牦朦蜩忄蝾疣, /耋;

v桉 耜 痤耱 桉镟疱龛 忸潲 镱忮瘐眍耱 耋耧屙玷, /耋.

锐觌桁 赅觇 猁 蝾 龛 猁腩 忡噼祛溴轳蜮, 镳 忸 箪屙屙棹 忮腓麒睇 S0 玎 聍蛤 钺疣珙忄 龛 礤疣耱忸痂禧 铖噤觐, 汔珙 镳. 亦赕 狍溴 镱豚汔螯, 黩 镳 镥疱戾忄龛 耋耧屙玷 扈牮钼 漕痤耠彘 沭囗桷 疣耱忸痂祛耱 NaCl 耋耧屙玷 漕 耱囹铟眍 忮腓觇 膻猁 忄痂圉梃 箐咫铋 耜铕铖蜩 镳铗铌 玎忮漕祛 礤 镳桠邃篁 恹镟溴龛 NaCl 铖 漕.

青觐 耦躔囗屙 钺爨 爨耨 镳邃耱噔弪 耠邃簋 忤溴:

dV镳 = dV恹 + dV桉 (2.10) dm镳 = dm0 + dm恹 沅 dV镳 = v镳 dt, dV桉 = v桉 dt, dm镳 = v镳 Sdt, dm0 = W dS0, dm恹 = S0 dV恹.

(2.11) 悟牦溧 dV恹 = v v 镳 桉 dt (2.12) v镳 S = W dS0 + S0 dV恹.

dt dt i i i i i i i i 蔫磬扈赅 觐眦屙蝠圉梃 NaCl 牦朦蝮疣朦眍 耩邃 绣 耔耱屐 (2.12), dS v镳 S = W + (v镳 v桉 ) S0 ;

dt dS W = v镳 S (v镳 v桉 ) S0 ;

dt v镳 v桉 篦蛤铎 镳 =, 桉 =, 桁邋:

W W dS (2.13) = S 镳 (镳 桉 ) S0.

dt 朽玟咫桠 镥疱戾眄 玎溧 磬鬣朦睇 篑腩忤, 祛 眍 磬轵 桧蝈沭嚯铄 恹疣驽龛 潆 滂磬扈觇 觐 鲥眚疣鲨 NaCl 牦朦蜩忄蝾疱:

dS = dt;

镳 S (镳 桉 )S S0 t dS = dt;

镳 S (镳 桉 )S t S 1 镳 S (镳 桉 )S ln = (t t ), 镳 桉 镳 S (镳 桉 )S 沅 S 觐眦屙蝠圉 NaCl 磬鬣朦睇 祛戾眚 怵 戾龛 t.

入:

镳 S (镳 桉 )S = e(镳 桉 )(tt ) ;

镳 S (镳 桉 )S i i i i i i i i 84 茵屙赍眸 . ., 缅忸疸桤 . ., 令痤怅钼 . .

镳 S(镳 桉 )S0 = = [镳 S (镳 桉 )S ] e(镳 桉 )(tt ) ;

镳 S0 (t) = S 镳 桉 (2.14) 镳 (镳 桉 )(tt ) S S e.

镳 桉 洛腓麒磬 桉 溧眄铎 恹疣驽龛 祛驽 镳桧 爨螯 赅 镱腩骅蝈朦睇, 蜞 铗痂鲟蝈朦睇 珥圜 龛. 悟痂鲟蝈朦眍 珥圜屙桢 桉 镳桧桁噱, 镳 戾痼, 镳 疱泱痦铎 疣耧疱耥屙梃 耋耧屙玷 铖噤赅 扈 镳 恹疣忄龛 D. salina 镳箐圊 镱 铗牮 礤犷. 奏耱睇 耠篦嚅: 镳 桉 = 0 趔黻鲨 (2.14) 镳孱狃噻钼噱蝰 忤潴 S0 (t) = S (S S ) e镳 (tt ) (2.15) 泽黻鲨 (2.14) 桁邋 镳邃咫 镳 t :

镳 st lim S0 (t) = S0 = S.

镳 桉 t 央邃钼囹咫, 磬 镳嚓蜩赍 忮腓麒磬 S0 耦 怵屐 st 礤 狍溴 耱疱扈螯 礤觐蝾痤 忮腓麒礤 S0 (耢.

痂. 2.7) 漕耱桡礤 甯 镳 篑蜞眍怆屙梃 耱圉桀磬 眍泐 滂磬扈麇耜钽 疣忭钼羼 (镳梓胳, 疣忭钼羼桢  st 篑蝾轺桠). 悟戾蜩 蜞赕, 黩 忮腓麒磬 S0 龛赅 礤 玎忤耔 铗 磬鬣朦睇 篑腩忤 S t 铒疱溴弪 蝾朦觐 箐咫 耜铕铖扈 桉镟疱龛 镳铗铌 耩邃.

i i i i i i i i 蔫磬扈赅 觐眦屙蝠圉梃 NaCl 牦朦蝮疣朦眍 耩邃 需. 2.7. 义铕弪梓羼觇 牮桠 滂磬扈觇 觐眦屙蝠 鲨 NaCl 牦朦蜩忄蝾疱 铗牮钽 蜩镟 (徉耨彘礤) 篑腩忤 噌耦膻蝽 礤镳屦眍 牦朦蝮瘥 (蹂祛耱囹) 镳 疣珉梓睇 磬鬣朦睇 篑腩忤 S, t (耢. 恹疣 驽龛 (2.14)), S = 100, = 0,1 桉 = 0,005.

i i i i i i i i 86 茵屙赍眸 . ., 缅忸疸桤 . ., 令痤怅钼 . .

殊噻桧屣疱瘥忭 疱骅 牦朦蜩忤痤忄龛. 篑腩忤 镳铎脲眄钽 牦朦蜩忤痤忄龛 扈牮钼钿铕铖脲 磬 徉珏 蝈镫梓睇 躅轳蜮 桉镱朦珙忄龛屐 羼蝈 耱忮眄钽 铖忮龛 蝾 桦 桧铋 戾疱 礤桤徨 睇 耱囗钼栩 镥痂钿梓眍耱 怦邈 蝈蹴铍钽梓羼觐泐 镳铞羼襦. 蜞觐 耔蝮圉梃 桉觌栩咫 怆弪 戾蝾 赈噻桧屣疱瘥忭铋 牦朦蝮瘥, 觐蝾瘥, 钿眍 耱铕铐, 珥圜栩咫铋 戾疱 箫痤弪 忮顸 蝈蹴铍 汨麇耜栝 镳铞羼 镳铊玮钿耱忄 徼铎囫覃, 漯筱铋 疣聒桊弪 忸珈铈眍耱 箫疣怆屙 觇礤蜩觐 痤耱 徼铖桧蝈珙 扈牮钼钿铕铖脲. 念耱囹铟眍 汨彡 镳 怆弪 皴 铗 戾蝾 觐眚痤脲 箫疣怆屙梃 忮腓 麒眍 觐眦屙蝠圉梃 疣耱忸鸶眄 耦脲, 鬣耱眍耱 NaCl. 如 囗嚯桤 礤镳屦眍 牦朦蝮瘥 耥, 黩 镳 桉镟疱龛 忸潲 镱忮瘐眍耱 耋耧屙玷 赅驿 镱 耠邃簋 镳铞邃箴 钺戾磬 狍溴 镱恹螯  忮 腓麒眢 漕 蝈 镱, 镱赅 铐 礤 漕耱桡礤 礤觐蝾痤 st 镱耱眄铋 Sn.

袜殇屐 蝈牦 忮腓麒眢 觐眦屙蝠圉梃 NaCl 镱 耠 镥疴铋 镳铞邃箴 钺戾磬 S1. 闰嚓, 玎 怵屐 t 戾驿 镳铞邃箴囔 钺戾磬 礤觐蝾瘥 钺 忸潲 w桉 桉镟痂蝰. 昨钺 耦躔囗栩 镱耱眈蜮 箐咫铋 耜 痤耱 镳铗铌 镳, 礤钺躅滂祛 耠栩 钺 耋耧屙玷 c 篦蛤铎 桉镟疱龛, . . 镱耠 耠桠 牦朦蜩忄蝾疱 漕腈屙 铖蜞螯 钺 Wc = W (w w桉 ). (2.16) 央邃钼囹咫, 祛骓 玎镨襦螯 耠邃簋 忮腓麒睇 i i i i i i i i 蔫磬扈赅 觐眦屙蝠圉梃 NaCl 牦朦蝮疣朦眍 耩邃 潆 觐眦屙蝠圉栝 NaCl 漕 镱耠 钺戾磬:

m0 W S0 =, S0 = S0, 桉 W w桉 W Wc S0 +(W Wc ) S W c S0 Wc 桉 S1 = = + 1 S, W w桉 W W 沅 S0 蝈牦 觐眦屙蝠圉 漕 钺戾磬 徨 桉镟疱 龛;

S0 蝈牦 觐眦屙蝠圉 漕 钺戾磬 篦蛤铎 桉 桉镟疱龛;

S1 蝈牦 觐眦屙蝠圉 镱耠 钺戾磬 徨 桉镟疱龛.

理嚯钽梓眍 恹疣驽龛 玎镨 潆 忮腓麒睇 S2, . . 镱耠 怛铕铋 镳铞邃箴 钺戾磬 Wc Wc S2 = S1 + 1 S = AS1 + B, W w桉 W 沅 Wc Wc (2.17) A=, B= 1 S.

W w桉 W 项漶蜞怆 恹疣驽龛 潆 S1 恹疣驽龛 潆 S2, 镱塍麒:

S2 = A(AS0 + B) + B.

溧脲, 潆 膻犷泐 眍戾疣 镳铞邃箴 钺戾磬 n 耧疣 忮潆桠 Sn = ASn1 +B = An S0 +B (1+A+A2 +...+An1 ).

i i i i i i i i 88 茵屙赍眸 . ., 缅忸疸桤 . ., 令痤怅钼 . .

湾蝠箐眍 玎戾蜩螯, 黩 禧 桁邋 溴腩 疱牦瘃桢 耋祆铋 沐铎弪痂麇耜铋 镳钽疱耨梃. 悟牦溧 铌铐鬣 蝈朦眍 玎镨 Sn 潆 膻犷泐 n.

An n (2.18) Sn = A S0 + B, A 1.

A 玉蝠屐 n 徨耜铐鬻眍耱, 镱塍麒 镳邃咫:

B st (2.19) lim Sn = Sn =.

1A n 央邃钼囹咫, 赅 耠篦噱 噌耦膻蝽 礤镳屦 睇 镳铗铌铎 耩邃, 忮腓麒磬 Sn 耦 怵屐屙屐 狍溴 st 耱疱扈螯 礤觐蝾痤 忮腓麒礤 Sn 漕耱桡礤 甯 镳 篑蜞眍怆屙梃 耱圉桀磬痦钽 滂磬扈麇耜钽 疣忭钼羼.

袜 痂耋黻 2.8 镳邃耱噔脲睇 蝈铕弪梓羼觇 牮桠 滂 磬扈觇 觐眦屙蝠圉梃 NaCl 耩邃 镳 恹疣忄龛 D. salina 赈噻桧屣疱瘥忭 戾蝾漕.

项 铒疱溴脲龛 [30], 潆 赈噻桧屣疱瘥忭铋 牦朦 蝮瘥 箐咫 耜铕铖螯 镳铗铌 镨蜞蝈朦眍 耩邃 麇疱 牦朦蜩忄蝾 镳邃耱噔弪 赅 1w (2.20) =, t W 沅 w 钺 玎戾睇 耋耧屙玷 镨蜞蝈朦眍 耩邃铋 玎 镳铎彐篁铌 t;

W 疣犷麒 钺 耋耧屙玷.

理嚯钽梓眍 恹疣驽龛 祛骓 玎镨襦螯 潆 箐咫 眍 耜铕铖蜩 桉镟疱龛:

1 w桉 (2.21) 桉 =.

t W i i i i i i i i 蔫磬扈赅 觐眦屙蝠圉梃 NaCl 牦朦蝮疣朦眍 耩邃 需. 2.8. 义铕弪梓羼觇 牮桠 滂磬扈觇 觐眦屙蝠 鲨 NaCl 牦朦蜩忄蝾疱 铗牮钽 蜩镟 (徉耨彘礤) 赈噻桧屣疱瘥忭铋 牦朦蝮疱 镳 疣珉梓睇 磬鬣朦 睇 篑腩忤 S, t (耢. 恹疣驽龛 (2.18), (2.23)), S = 100, = 0,1, 桉 = 0,005.

i i i i i i i i 90 茵屙赍眸 . ., 缅忸疸桤 . ., 令痤怅钼 . .

沅 w桉 钺 桉镟疱龛 玎 怵屐 t.

蔓疣驽龛 (2.18) 祛骓 玎镨襦螯 耠邃簋 钺 疣珙 An B B n Sn = A S0 + A1 A 桦, 篦蛤铎 (2.19) Sn = An (S0 Sn ) + Sn.

st st (2.22) 泥脲, 篦蛤铎 (2.20) (2.21) 桤 (2.16) 耠邃箦 w w桉 Wc =1 = 1 t ( 桉 ).

W W 悟颃溧 Wc Wc 1 t ( 桉 ) A= = =.

W w桉 W W t 桉 1 t 桉 项漶蜞怆 镱塍麇眄 疱珞朦蜞 (2.22), 铌铐鬣蝈朦 眍 桁邋 n 1 t ( 桉 ) st st (S0 Sn ) + Sn. (2.23) Sn = 1 t 桉 闰嚓, 蝈铕弪梓羼觇 囗嚯桤 镱赅玎, 黩 赅 耠篦噱 噌耦膻蝽 礤镳屦眍, 蜞 赈噻桧屣疱瘥 眍 牦朦蝮疱 Dunaliella salina 镳 镱耱眈蜮 觐眦屙 蝠圉梃 NaCl 镱溧忄屐铋 牦朦蜩忄蝾 镨蜞蝈朦眍 耩邃, 觐眦屙蝠圉 NaCl 牦朦蝮疣朦眍 耩邃 狍溴 i i i i i i i i 蔫磬扈赅 觐眦屙蝠圉梃 NaCl 牦朦蝮疣朦眍 耩邃 耱疱扈螯 礤觐蝾痤 镱耱眄铋 S st. 亦赕 囗嚯桤 疣琊噌铗囗眍 祛溴腓 镱赅玎, 黩 珥囗桢 忮腓麒睇 箐咫铋 耜铕铖蜩 桉镟疱龛 忸潲 镱忮瘐眍耱 徉 皴轫 镱玮铍栩 脲汴 漕徼螯 镱滗屦驵龛 玎溧眄钽 珥圜屙 耦敫眍耱 牦朦蝮疣朦眍 耩邃 镱祛 桤戾礤龛 忮腓麒睇 箐咫铋 耜铕铖蜩 镳铗铌 镨蜞 蝈朦眍 耩邃.

i i i i i i i i 青觌屙桢 语疣怆屙桢 镳铞羼耦 牦朦蜩忤痤忄龛 Dunaliella salina 镳铎脲眄 爨聒蜞徉 鲥朦 漕耱桄 龛 爨犟桁嚯铋 镳钿箨蜩忭铖蜩 镱 徼铎囫皴 桦 镱 赅觐祗-腓犷 徼铛桁梓羼觐祗 镱赅玎蝈膻 徼铎囫覃 (磬镳桁屦, -赅痤蜩眢) 镱 镳彐礤祗 铖蜞蛤 漕耱 蝾黜 耠铈眍 玎溧麇, 礤耢铗 磬 祉钽铍弪龛 桉 耠邃钼囗 溧眄铋 钺豚耱. 袜 铒 镱赅琨忄 弪, 黩 磬 镳铊玮钿耱忮 铖钺铄 忭桁囗桢 礤钺躅滂祛 箐咫栩 耠邃簋 忸镳铖囔: 1. 蔓犷痼 耦铗忮蝰怏 泐 蝠遽钼囗 镳铊玮钿耱忄 囔爨 D. salina, 磬 镳桁屦, 潆 镳钿箨鲨 -赅痤蜩磬 磬 铖眍忮 疱珞朦 蜞蝾 镳钼邃疙睇 耧屦桁弪钼 祛骓 篁忮疰溧螯, 黩 磬栳铍邋 镱漉钿 怆弪 IBSS-2 桤 觐腚尻 鲨 肃眈觐 (软赁). 2. 蔓犷痼 镨蜞蝈朦眍 耩 潲 潆 牦朦蜩忤痤忄龛. 袜栳铍邋 铒蜩爨朦睇 忄 痂囗蝾 怆弪 祛滂翳鲨痤忄眄1 耩邃 Ben-Amotz (耢. 耱. 50). 3. 项滗屦驵龛 嚯铍钽梓羼觐 麒耱 螓 牦朦蝮瘥 D. salina. 碾 钽 耠邃箦 觐眦屙蝠圉棹 NaCl 耩邃 怦邈溧 镱滗屦骅忄螯 磬 箴钼礤 120 /. , 磬觐礤. 4. 务钺铄 忭桁囗桢 耠邃箦 箐咫栩 恹犷痼 填滂翳赅鲨 玎觌噱蝰 漕徉怆屙梃 120 / NaCl 忪羼蝾 60.

i i i i i i i i 青觌屙桢 戾蝾溧 牦朦蜩忤痤忄龛. 碾 D. salina 磬栳铍邋 镳 屐脲禧 铌噻嚯 戾蝾 礤镳铒铕鲨铐嚯-镳铗铟眍 牦朦蝮瘥, 觐蝾瘥 镱玮铍弪 疱螯 鲥臌 玎溧 蝈蹴铍钽梓羼觐泐 踵疣牝屦.

i i i i i i i i 扬桉铌 腓蝈疣蝮瘥 1. 泪潴腚噱 . ., 彦戾礤黻 . . 软蝈眈桠磬 牦朦蝮疣 Dunaliella salina Teod. 礤觐蝾痂 甯 翳 玷铍钽梓羼觇 踵疣牝屦桉蜩觇 // 澡玷铍钽 疣 蝈龛. 1974. . 21. . 11451153.

2. 拎觌囗钼 . ., 嘴桦屙觐 . . 项塍麇龛 -赅 痤蜩磬 (钺珙) // 蓐铗艴眍腩汨 疱耋瘃铖徨疱驽 龛. 2000. 4. . 3235.

3. 铃龛 . ., 谚潼觐 . ., 茵屙赍眸 . .

蓓屦沐蜩赅 纛蝾耔眚彗桊簋 牦朦蝮瘥 扈牮钼 漕痤耠彘 / 项 疱. . . 暑怵钼. 皖忸耔徼瘃:

袜箨. 盐, 1980. 126 .

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镳 疣珉梓龛 篑腩忤 牦朦蜩忤痤忄龛 // 离 腩汨. 2003. . 13, 2. . 137147.

5. 码噤桁桊钼 . . 与疣耱痼牝箴磬 铕汔龛玎鲨 觌弪觇 Dunaliella salina 甯 趔黻鲨铐嚯桢 桤 戾礤龛 玎忤耔祛耱 铗 桧蝈眈桠眍耱 疋弪 i i i i i i i i 扬桉铌 腓蝈疣蝮瘥 蝈祜屦囹箴 // 澡玷铍钽 疣耱屙栝. 1978. . 25. . 571576.

6. 码噤桁桊钼 . ., 彦戾礤黻 . . 软蝈眈桠磬 牦朦蝮疣 钿眍觌弪铟睇 忸漕痤耠彘 (桧耱痼牿 镱 镥疴梓睇 桉稃蜞龛, 恹溴屐 桤 镳桊钿 皴脲牿桀龛痼屐 纛痨 纛蝾噔蝾蝠铘睇 钿眍觌 蝾黜 忸漕痤耠彘) / 项 疱. . . 丸麒镱痤忤 鬣. .: 劳 蜒研, 1962. 60 .

7. 缅忸疸桤 . ., 令痤怅钼 . . 蔫磬扈赅 徼铎囫覃 Dunaliella salina 篑腩忤 礤镳屦眍泐 牦朦蜩 忤痤忄龛 // 蓐铍钽 祛. 2005. . 67. . 3537.

8. 蔫忄忤 . ., 暑稃蝾 . . 蔫磬扈赅 桤 戾礤龛 徼铛桁梓羼觐泐 耦耱噔 铗溴朦睇 铕汔 眍 扈滂 Mytilus galoprovincialis 镳 桧蝾犟桕 鲨 耦痤 // 蓐铍钽 祛. 1988. . 26. . 8086.

9. 酿铌钼 I. ., 鼠珥弼钼 . ., 项镱忄 . . 蚂i耱 赅痤蜩眢 忸漕痤耱i Dunaliella salina Teod. 镳 忤 痤忄眄i 磬 疣飅 豚犷疣蝾痦桴 箪钼圊 // 睦 有研. 彦. . 1967. . 8. . 736739.

10. 枢蜞腩 牦朦蝮 扈牮钼钿铕铖脲 觐腚尻鲨 蜒研. : 仍 欣, 1991. 226 .

11. 翳玷铍钽梓羼觐 踵疣牝屦桉蜩赍 裩lorella sp. 镳 恹耦觇 耱疱爨朦龛 蝈祜屦囹箴圊. II. 如戾礤龛 i i i i i i i i 96 茵屙赍眸 . ., 缅忸疸桤 . ., 令痤怅钼 . .

徼铖桧蝈玎, 箅疣耱痼牝箴 嚓蜩忭铖蜩 纛蝾 耔眚弪梓羼觐泐 囡镟疣蜞 蹼铕咫臌 镳 疣珙狴屙梃 觌弪铟睇 趔黻鲨 耱疱爨朦眍 蝈祜屦囹箴铋 / . . 彦戾礤黻, . . 码噤桁桊钼, .. 勿脲 囗耜栝, 漯. // 澡玷铍钽 疣耱屙栝. 1969. . 16. . 210220.

12. 锑颃 . . 嘛腓 轭韎 Na, Mg, Cl 蜞 SO 磬 餴耱, 疣珈眍驽眄 蜞 赅痤蜩眍篁忸疱眄 忸漕 痤耱i Dunaliella salina Teod. // 雨. 犷. 骟痦. 1965. . 22, 5. . 39.

13. 锑颃 . . 锑耦忄 牦朦蝮疣 赅痤蜩眍眍耥羁 忸 漕痤耱i Dunaliella salina teod. // 雨. 犷. 骟痦. 1966. . 23, 2. . 1218.

14. 锑颃 . . 填痿铍钽, 耔耱屐囹桕, 铍钽, 沐钽疣翳麇耜铄 疣耧疱溴脲龛 痤溧 Dunaliella Teod.

镥瘃镥牝桠 镳嚓蜩麇耜钽 桉镱朦珙忄龛. .:

袜箨. 潴礻, 1973. 244 .

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i i i i i i i i 扬桉铌 腓蝈疣蝮瘥 17. 惕朦觐 . . 如篦屙桢 镱蝠遽眍耱彘 溻篚 忤漕 忸漕痤耠彘 扈礤疣朦龛 铕汔龛麇耜桴 觐祜 礤眚圊 耩邃 // 洛耱龛 堂. 1962. 1. . 1824.

18. 惕朦觐 . . 如篦屙桢 翳玷铍钽梃 镨沆屙蝾 钺疣珙忄龛 珏敫眍 忸漕痤耠 Dunaliella: 棱蝾疱.

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i i i i i i i i 毋豚怆屙桢 骡邃屙桢 1. Dunaliella salina Teod. 赅 钺牝 徼铗艴眍腩 汨 1.1. 清敫磬 扈牮钼钿铕铖朦 Dunaliella salina. 1.2. 务钺屙眍耱 桧蝈眈桠眍泐 牦朦蜩忤痤 忄龛 Dunaliella salina.......... 1.3. IBSS-2 镥瘃镥牝桠睇 囔 潆 镳铎脲眄钽 牦朦蜩忤痤忄龛.... 1.4. 蓄耱 Dunaliella salina 耢屮囗眍 牦朦蝮疱.................. 2. 物蜩扈玎鲨 镳铞羼耦 牦朦蜩忤痤忄龛 Dunaliella salina 2.1. 物蜩扈玎鲨 怵屐屙 怅膻麇龛 镳 蝾赅 忮腓麒睇 箐咫铋 耜铕铖蜩 镳 蝾赅.................... 2.2. 蔫磬扈赅 觐眦屙蝠圉梃 NaCl 牦朦 蝮疣朦眍 耩邃.............. 青觌屙桢 i i i i i i i i R. P. Trenkenshu, R. G. Gevorgiz, A. B. Borovkov The fundamentals of industrial cultivation of Dunaliella Teod. Sevastopol: ECOSI朒ydrophisica, salina 2005. 103 p.

The results of investigations of Dunaliella salina Teod. production characteristics provided at industrial conditions have been presented in this book.

The attention has been shared out to the questions of cultivation processes optimization, biology and biochemistry of green microalgae D. salina.

Monograph presents interest for the biology specialists also it will be useful for the postgraduate students and students.

茵屙赍眸 . ., 缅忸疸桤 . ., 令痤怅钼 . .

务眍忤 镳铎桉腩忸泐 牦朦蜩怏忄眄 捏磬雐咫腓 耦腩眍忸漤铋 (Dunaliella salina Teod.). 彦忄耱铒铍:

菔窝I柮i漯铘i玷赅, 2005. 103 .

祛眍沭圄i 镳邃耱噔脲眍 疱珞朦蜞蜩 漕耠i滏屙 镳钿箨鰅轫桴 踵疣牝 痂耱桕 Dunaliella salina Teod., 阨 镳钼钿桦桉 忤痤犴梓桴 箪钼圊. 橡桎i 脲眍 筲嚆 镨蜞眄 铒蜩靑玎鰅 镳铞羼 牦朦蜩怏忄眄, 醝铍钽i 蜞 醝铛i靑 珏敫眍 靑牮钼钿铕铖騣 D. salina.

填眍沭圄i 耱囗钼栩 i眚屦羼 潆 羿蹊怫i-醝铗艴眍腩鉯. 亦觐 狍溴 觐痂耥 囫飅疣眚囔 鈏滹钼i漤桴 耧弼i嚯铖蝈 i 耱箐屙蜞 忤 磬怊嚯 龛 玎觌噤i.

i i i i i i i i 畜漕朦 相怆钼梓 茵屙赍眸, 玎忮潴栝 铗溴腩 徼铗艴眍腩汨 翳蝾疱耋瘃钼.

缅忸疸桤 畜耠囗 缅铕汨邂梓, 磬篦睇 耦蝠箐龛 铗溴豚 徼铗艴眍腩汨 翳蝾疱耋瘃钼.

令痤怅钼 理漯彘 令痂耦忤, 祀噤 磬篦睇 耦蝠箐龛 铗溴豚 徼铗艴眍腩汨 翳蝾疱耋瘃钼 软耱栩篁 徼铍钽梃 睇 祛疱 桁. . . 暑忄脲怦觐泐 镳. 袜蹊祛忄, 2, . 彦忄耱铒铍, 署 E-mail: trenkens@yandex.ru http://biotex.ibss.org.ua 义脲纛: (+038)(0692)55-07- 勿桡桧嚯-爨赍 镱溷铗钼桦 . . 缅忸疸桤 镟赍蝈 LTEX 2 桉 A 镱朦珙忄龛屐 觇痂腚梓羼觇 梏蝾 . 隋徨溴忄 PSCyr (v0.4c).

唁囗 磬犷 30.11.2005. 项滹桉囗 镥鬣蜩 27.12. 襄鬣螯 铘皴蝽. 灶痨囹 70108 1 /16. 绵痦栩箴 嚓噤屐梓羼赅.

吾: 镥鬣蝽 腓耱钼. 喻.-桤... 诣疣 . 青赅.

拖 饰讶-描漯铘桤桕嗷, 99005 彦忄耱铒铍, 箅. 隋龛磬, 28. 砚桎 蝈朦耱忸 泐耋溧瘃蜮屙眍 疱汨耱疣鲨: 皴痂 氖 914 铗 16.02.02 .

i i i i

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